Type locality: "Small shallow mountain creek branch, 55 kilometers from the Mimongo-Mbigou Road, near Iméno Mbila, Du Chaillu Massif, southern Gabon. Holotype: MRAC 80-54-P-1536. Paratypes: (52) MRAC 80-54-P-1537-45 (9), 80-54-P-1546-85 (40)"