Wolfgang Eberl wrote a special about this species group (DKG Journal 2016-1) I agree what he wrote: bochtleri and herzogi are confined to central Gabon (red oval). All others are related to herzogi. Some of them with very aberrant phenotypes and DNA. Complex group! - JVDZ

Ayene GEMLBJ 03-30 Edoum Akurenam GEMGSL 19-13 Efulan GEGVPO 18-12 GEMLH 2016-27

Mbam GEMHS 00-12 Medouneu JH-52 Sork Oveng GEMLCG 07-23 Oveng is about halfway on the (N2) road between Mitzic and Lalara in northern Gabon. Zomoko GBG 92-25 Type locality: Fast running brook about 3 kilometers north of Zoumoukou or 16 kilometers north of Lalara on the road to Mitzic and Oyem, northern Gabon, 0°27'N, 12°15'E.